The western limb of the Hellenic Arc defines the boundary of a large intracontinental active extensional domain covering the Aegean and the southwestern part of the Balkan peninsula. Along this boundary a transition from collision in the north to subduction in the south is associated with post-Miocene clockwise rotations of ∼50 • . We present a new GPS velocity field that, with new permanent station velocities in Albania, Bulgaria, Kosovo, Montenegro, and Northern Macedonia, provides insights into previously unresolved aspects on the large-scale dynamics of continental lithosphere and on the relation between long and short-term kinematics. In particular we address (1) the kinematic description of the collision/subduction transition, (2) the relation between long-term finite rotations with geodeticallymeasured instantaneous rotations, (3) the forces maintaining and resisting the deformation of the extensional domain and (4) the extent of its northern boundary. We use the analogy of rigid elongate inclusions in the velocity field to predict the senses and approximate rates of rotation of crustal blocks in the deforming continental region. Across the collision/subduction transition zone a large rotational pattern in the velocity field is found whose spatial pattern and sense of rotation is compatible with the observed paleomagnetic rotations through the occurrence of fault-bounded block rotations. Geodetic observations show that clockwise vertical axis rotation of the western limb of the Hellenic margin is active today, with distribution and rates that are essentially the same as that defined by the paleomagnetic data for the last 5 Ma. The maximum rates of the rotational component of the velocity field is found along the wide arrangement of fault-bounded blocks in central Greece that accomodate the NE-SW dextral shear transferred from the northern Aegan trough. The northern limit of the rotating margin defines a hinge (Scutari-Pec transverse zone) that has remained stationary throughout the Middle Miocene (relative to the upper plate of the collision/subduction system) with important implications for the distribution of strength of the western margin of the Hellenic Arc. The distribution and style of deformation along the western boundary of the extensional domain is controlled by the relation between gravitational forces, driving the flow of crustal material towards the low-lying adjacent regions sea floor, and heterogeneous resistive forces along the collision/subduction boundary.