Abstract “Stone runs” is the Falklands vernacular term for openwork boulder accumulations, which include extensive blockstreams like the famous Darwin “stone-river” and associated features such as stone stripes. Since the early 20th century, they have been interpreted as the product of a suite of periglacial processes, including frost-wedging, gelifluction, frost heave, frost-sorting and snowmelt runoff. Following a literature review, the results of recent field investigations of the valley-floor blockstreams of East Falkland are presented. Access to the internal structure of these forms provides evidence for the existence of a three-fold profile, with clear vertical size gradation presenting striking similarities with an inverted weathering profile. Micromorphological analyses, SEM, XRD, thin sections and grain-size analyses lead to the hypothesis of an alternative model of stone run formation. It is suggested that the material forming the stone run profile lato sensu (including the superficial pavement) is not of periglacial origin, but derives directly from the stripping and accumulation downslope of a regolith, possibly Tertiary in age and formed under subtropical or temperate conditions. The valley-floor stone runs should, therefore, be considered as complex polygenetic landforms that may have formed according to a six-stage scenario, including in situ chemical weathering, regolith stripping by mass movements, soil formation, further regolith stripping, downslope accumulation and matrix washing-out (all phases possibly achieved by the Early Quaternary). Periglacial reworking of the stone run material would have operated at a “final” stage, i.e. during Quaternary cold stages, with boulder bioweathering and limonite-staining operating during the temperate intervals including the present one. The suggested antiquity of the Falklands blockstreams is in accordance with Caine's pioneer interpretation of Tasmania blockfields and with recent analyses and cosmogenic datings of blockfields from Scandinavia and North America.