Epigenetic control is involved in molecular dialogue in plantmicrobe symbiosis Epigenetic processes affect gene expression without changes in DNA sequences. In eukaryotes, the most common epigenetic modification is a DNA cytosine methylation consisting of the addition of a methyl group (CH 3) to a cytosine nucleotide. In plants, genomic DNA (gDNA) methylation is an extensively studied epigenetic mark that can occur in three sequence contexts, namely at the sites CG, CHG, and CHH (where H is A, T, C). Moreover, it is known that transposable elements (TEs) are presumed to be densely methylated and are thus inactive. In recent years, molecular mechanisms of DNA methylation in genetic model plant species have been extensively studied (To & Kakutani, 2022; Kakoulidou & Johannes, 2023).