<?xml version="1.0" encoding="utf-8"?>
<TEI xmlns="http://www.tei-c.org/ns/1.0" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xmlns:hal="http://hal.archives-ouvertes.fr/" xmlns:gml="http://www.opengis.net/gml/3.3/" xmlns:gmlce="http://www.opengis.net/gml/3.3/ce" version="1.1" xsi:schemaLocation="http://www.tei-c.org/ns/1.0 http://api.archives-ouvertes.fr/documents/aofr-sword.xsd">
  <teiHeader>
    <fileDesc>
      <titleStmt>
        <title>HAL TEI export of hal-04036146</title>
      </titleStmt>
      <publicationStmt>
        <distributor>CCSD</distributor>
        <availability status="restricted">
          <licence target="https://creativecommons.org/publicdomain/zero/1.0/">CC0 1.0 - Universal</licence>
        </availability>
        <date when="2026-05-24T00:45:01+02:00"/>
      </publicationStmt>
      <sourceDesc>
        <p part="N">HAL API Platform</p>
      </sourceDesc>
    </fileDesc>
  </teiHeader>
  <text>
    <body>
      <listBibl>
        <biblFull>
          <titleStmt>
            <title xml:lang="en">Zonkey : a simple, accurate and sensitive pipeline to genetically identify equine F1-hybrids in archaeological assemblages</title>
            <author role="aut">
              <persName>
                <forename type="first">Mikkel</forename>
                <surname>Schubert</surname>
              </persName>
              <idno type="halauthorid">983067-0</idno>
              <affiliation ref="#struct-3549"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Marjan</forename>
                <surname>Mashkour</surname>
              </persName>
              <email type="md5">cfc780eb95568693df99840369534565</email>
              <email type="domain">mnhn.fr</email>
              <idno type="idhal" notation="string">marjan-mashkour</idno>
              <idno type="idhal" notation="numeric">176173</idno>
              <idno type="halauthorid" notation="string">6741-176173</idno>
              <idno type="ORCID">https://orcid.org/0000-0003-3630-9459</idno>
              <idno type="IDREF">https://www.idref.fr/057725977</idno>
              <affiliation ref="#struct-1225933"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Charleen</forename>
                <surname>Gaunitz</surname>
              </persName>
              <email type="md5">e387650331509191765fdfebfa620dc9</email>
              <email type="domain">sund.ku.dk</email>
              <idno type="idhal" notation="numeric">1532017</idno>
              <idno type="halauthorid" notation="string">1298927-1532017</idno>
              <idno type="ORCID">https://orcid.org/0000-0003-3681-528X</idno>
              <affiliation ref="#struct-3549"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Antoine</forename>
                <surname>Fages</surname>
              </persName>
              <email type="md5">caf0258fcb7996126cd7559ad891eb58</email>
              <email type="domain">gmail.com</email>
              <idno type="idhal" notation="numeric">1178454</idno>
              <idno type="halauthorid" notation="string">1298928-1178454</idno>
              <idno type="ORCID">https://orcid.org/0000-0001-6985-4360</idno>
              <idno type="GOOGLE SCHOLAR">https://scholar.google.fr/citations?user=w7aPYDAAAAAJ</idno>
              <idno type="IDREF">https://www.idref.fr/24181300X</idno>
              <idno type="VIAF">https://viaf.org/viaf/94158125211214932902</idno>
              <affiliation ref="#struct-445683"/>
              <affiliation ref="#struct-3549"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Andaine</forename>
                <surname>Seguin-Orlando</surname>
              </persName>
              <email type="md5">023067d52ad7f9bdf75d790c74df2605</email>
              <email type="domain">univ-tlse3.fr</email>
              <idno type="idhal" notation="string">andaine-seguin-orlando</idno>
              <idno type="idhal" notation="numeric">776447</idno>
              <idno type="halauthorid" notation="string">983072-776447</idno>
              <idno type="ORCID">https://orcid.org/0000-0002-8265-3229</idno>
              <idno type="IDREF">https://www.idref.fr/110945514</idno>
              <affiliation ref="#struct-3549"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Shiva</forename>
                <surname>Sheikhi</surname>
              </persName>
              <idno type="halauthorid">1621920-0</idno>
              <affiliation ref="#struct-1225933"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Ahmed</forename>
                <surname>Alfarhan</surname>
              </persName>
              <idno type="halauthorid">2318756-0</idno>
              <affiliation ref="#struct-302265"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Saleh</forename>
                <surname>Alquraishi</surname>
              </persName>
              <idno type="halauthorid">1298933-0</idno>
              <affiliation ref="#struct-302265"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Khaled A.S.</forename>
                <surname>Al-Rasheid</surname>
              </persName>
              <idno type="halauthorid">1610015-0</idno>
              <affiliation ref="#struct-302265"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Richard</forename>
                <surname>Chuang</surname>
              </persName>
              <idno type="halauthorid">2135820-0</idno>
              <affiliation ref="#struct-300666"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Luca</forename>
                <surname>Ermini</surname>
              </persName>
              <idno type="halauthorid">983066-0</idno>
              <affiliation ref="#struct-3549"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Cristina</forename>
                <surname>Gamba</surname>
              </persName>
              <idno type="halauthorid">983071-0</idno>
              <affiliation ref="#struct-3549"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Jaco</forename>
                <surname>Weinstock</surname>
              </persName>
              <idno type="halauthorid">1610074-0</idno>
              <affiliation ref="#struct-300666"/>
            </author>
            <author role="aut">
              <persName>
                <forename type="first">Onar</forename>
                <surname>Vedat</surname>
              </persName>
              <idno type="idhal" notation="numeric">1178464</idno>
              <idno type="halauthorid" notation="string">2761086-1178464</idno>
              <idno type="ORCID">https://orcid.org/0000-0002-8359-243X</idno>
              <idno type="RESEARCHERID">http://www.researcherid.com/rid/D-1582-2019</idno>
              <idno type="RESEARCHERID">http://www.researcherid.com/rid/http://www.researcherid.com/rid/D-1582-2019</idno>
              <affiliation ref="#struct-495165"/>
            </author>
            <author role="crp">
              <persName>
                <forename type="first">Ludovic</forename>
                <surname>Orlando</surname>
              </persName>
              <email type="md5">7585ec3f47399a860c08466925c6c7e0</email>
              <email type="domain">univ-tlse3.fr</email>
              <idno type="idhal" notation="string">ludovicorlando</idno>
              <idno type="idhal" notation="numeric">1083415</idno>
              <idno type="halauthorid" notation="string">312103-1083415</idno>
              <idno type="ORCID">https://orcid.org/0000-0003-3936-1850</idno>
              <idno type="IDREF">https://www.idref.fr/080106951</idno>
              <idno type="RESEARCHERID">http://www.researcherid.com/rid/A-8932-2013</idno>
              <idno type="RESEARCHERID">http://www.researcherid.com/rid/http://www.researcherid.com/rid/A-8932-2013</idno>
              <idno type="GOOGLE SCHOLAR">https://scholar.google.fr/citations?user=zSlox0oAAAAJ</idno>
              <idno type="VIAF">https://viaf.org/viaf/66751689</idno>
              <idno type="ISNI">http://isni.org/isni/0000000041962127</idno>
              <affiliation ref="#struct-445683"/>
              <affiliation ref="#struct-3549"/>
            </author>
            <editor role="depositor">
              <persName>
                <forename>Ludovic</forename>
                <surname>ORLANDO</surname>
              </persName>
              <email type="md5">7585ec3f47399a860c08466925c6c7e0</email>
              <email type="domain">univ-tlse3.fr</email>
            </editor>
          </titleStmt>
          <editionStmt>
            <edition n="v1" type="current">
              <date type="whenSubmitted">2024-11-13 17:40:31</date>
              <date type="whenModified">2025-10-22 18:04:09</date>
              <date type="whenReleased">2024-11-15 16:45:16</date>
              <date type="whenProduced">2017-02</date>
              <date type="whenEndEmbargoed">2024-11-13</date>
              <ref type="file" target="https://hal.science/hal-04036146v1/document">
                <date notBefore="2024-11-13"/>
              </ref>
              <ref type="file" subtype="author" n="1" target="https://hal.science/hal-04036146v1/file/__filestore.soton.ac.uk_users_jw20_mydocuments_e-papers%252520and%252520books_Zonkey_uncorrected_proof_JASC.pdf" id="file-4781429-4163524">
                <date notBefore="2024-11-13"/>
              </ref>
              <ref type="externalLink" target="https://eprints.soton.ac.uk/405270/1/__filestore.soton.ac.uk_users_jw20_mydocuments_e-papers%2520and%2520books_Zonkey_uncorrected_proof_JASC.pdf"/>
            </edition>
            <respStmt>
              <resp>contributor</resp>
              <name key="691610">
                <persName>
                  <forename>Ludovic</forename>
                  <surname>ORLANDO</surname>
                </persName>
                <email type="md5">7585ec3f47399a860c08466925c6c7e0</email>
                <email type="domain">univ-tlse3.fr</email>
              </name>
            </respStmt>
          </editionStmt>
          <publicationStmt>
            <distributor>CCSD</distributor>
            <idno type="halId">hal-04036146</idno>
            <idno type="halUri">https://hal.science/hal-04036146</idno>
            <idno type="halBibtex">schubert:hal-04036146</idno>
            <idno type="halRefHtml">&lt;i&gt;Journal of Archaeological Science&lt;/i&gt;, 2017, 78, pp.147-157. &lt;a target="_blank" href="https://dx.doi.org/10.1016/j.jas.2016.12.005"&gt;&amp;#x27E8;10.1016/j.jas.2016.12.005&amp;#x27E9;&lt;/a&gt;</idno>
            <idno type="halRef">Journal of Archaeological Science, 2017, 78, pp.147-157. &amp;#x27E8;10.1016/j.jas.2016.12.005&amp;#x27E9;</idno>
            <availability status="restricted">
              <licence target="https://about.hal.science/hal-authorisation-v1/">HAL Authorization<ref corresp="#file-4781429-4163524"/></licence>
            </availability>
          </publicationStmt>
          <seriesStmt>
            <idno type="stamp" n="SHS">Sciences de l'Homme et de la Société</idno>
            <idno type="stamp" n="MNHN">Muséum National d'Histoire Naturelle</idno>
            <idno type="stamp" n="UNIV-TLSE3">Université de Toulouse</idno>
            <idno type="stamp" n="CNRS">CNRS - Centre national de la recherche scientifique</idno>
            <idno type="stamp" n="FRANTIQ">FRANTIQ Fédération et ressources sur l’Antiquité</idno>
            <idno type="stamp" n="AASPE" corresp="MNHN">Archéozoologie et Archéobotanique - Sociétés, Pratiques et Environnements</idno>
            <idno type="stamp" n="ALLIANCE-SU"> Alliance Sorbonne Université</idno>
            <idno type="stamp" n="UNIV-UT3">Université Toulouse 3</idno>
            <idno type="stamp" n="UT3-INP">Université de Toulouse / Toulouse INP</idno>
            <idno type="stamp" n="UT3-TOULOUSEINP">Université de Toulouse / Toulouse INP</idno>
            <idno type="stamp" n="TEST3-HALCNRS">TEST3-HALCNRS</idno>
            <idno type="stamp" n="TEST4-HALCNRS">collection test</idno>
            <idno type="stamp" n="TEST5-HALCNRS">collection test 5</idno>
          </seriesStmt>
          <notesStmt>
            <note type="audience" n="2">International</note>
            <note type="popular" n="0">No</note>
            <note type="peer" n="1">Yes</note>
          </notesStmt>
          <sourceDesc>
            <biblStruct>
              <analytic>
                <title xml:lang="en">Zonkey : a simple, accurate and sensitive pipeline to genetically identify equine F1-hybrids in archaeological assemblages</title>
                <author role="aut">
                  <persName>
                    <forename type="first">Mikkel</forename>
                    <surname>Schubert</surname>
                  </persName>
                  <idno type="halauthorid">983067-0</idno>
                  <affiliation ref="#struct-3549"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Marjan</forename>
                    <surname>Mashkour</surname>
                  </persName>
                  <email type="md5">cfc780eb95568693df99840369534565</email>
                  <email type="domain">mnhn.fr</email>
                  <idno type="idhal" notation="string">marjan-mashkour</idno>
                  <idno type="idhal" notation="numeric">176173</idno>
                  <idno type="halauthorid" notation="string">6741-176173</idno>
                  <idno type="ORCID">https://orcid.org/0000-0003-3630-9459</idno>
                  <idno type="IDREF">https://www.idref.fr/057725977</idno>
                  <affiliation ref="#struct-1225933"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Charleen</forename>
                    <surname>Gaunitz</surname>
                  </persName>
                  <email type="md5">e387650331509191765fdfebfa620dc9</email>
                  <email type="domain">sund.ku.dk</email>
                  <idno type="idhal" notation="numeric">1532017</idno>
                  <idno type="halauthorid" notation="string">1298927-1532017</idno>
                  <idno type="ORCID">https://orcid.org/0000-0003-3681-528X</idno>
                  <affiliation ref="#struct-3549"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Antoine</forename>
                    <surname>Fages</surname>
                  </persName>
                  <email type="md5">caf0258fcb7996126cd7559ad891eb58</email>
                  <email type="domain">gmail.com</email>
                  <idno type="idhal" notation="numeric">1178454</idno>
                  <idno type="halauthorid" notation="string">1298928-1178454</idno>
                  <idno type="ORCID">https://orcid.org/0000-0001-6985-4360</idno>
                  <idno type="GOOGLE SCHOLAR">https://scholar.google.fr/citations?user=w7aPYDAAAAAJ</idno>
                  <idno type="IDREF">https://www.idref.fr/24181300X</idno>
                  <idno type="VIAF">https://viaf.org/viaf/94158125211214932902</idno>
                  <affiliation ref="#struct-445683"/>
                  <affiliation ref="#struct-3549"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Andaine</forename>
                    <surname>Seguin-Orlando</surname>
                  </persName>
                  <email type="md5">023067d52ad7f9bdf75d790c74df2605</email>
                  <email type="domain">univ-tlse3.fr</email>
                  <idno type="idhal" notation="string">andaine-seguin-orlando</idno>
                  <idno type="idhal" notation="numeric">776447</idno>
                  <idno type="halauthorid" notation="string">983072-776447</idno>
                  <idno type="ORCID">https://orcid.org/0000-0002-8265-3229</idno>
                  <idno type="IDREF">https://www.idref.fr/110945514</idno>
                  <affiliation ref="#struct-3549"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Shiva</forename>
                    <surname>Sheikhi</surname>
                  </persName>
                  <idno type="halauthorid">1621920-0</idno>
                  <affiliation ref="#struct-1225933"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Ahmed</forename>
                    <surname>Alfarhan</surname>
                  </persName>
                  <idno type="halauthorid">2318756-0</idno>
                  <affiliation ref="#struct-302265"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Saleh</forename>
                    <surname>Alquraishi</surname>
                  </persName>
                  <idno type="halauthorid">1298933-0</idno>
                  <affiliation ref="#struct-302265"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Khaled A.S.</forename>
                    <surname>Al-Rasheid</surname>
                  </persName>
                  <idno type="halauthorid">1610015-0</idno>
                  <affiliation ref="#struct-302265"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Richard</forename>
                    <surname>Chuang</surname>
                  </persName>
                  <idno type="halauthorid">2135820-0</idno>
                  <affiliation ref="#struct-300666"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Luca</forename>
                    <surname>Ermini</surname>
                  </persName>
                  <idno type="halauthorid">983066-0</idno>
                  <affiliation ref="#struct-3549"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Cristina</forename>
                    <surname>Gamba</surname>
                  </persName>
                  <idno type="halauthorid">983071-0</idno>
                  <affiliation ref="#struct-3549"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Jaco</forename>
                    <surname>Weinstock</surname>
                  </persName>
                  <idno type="halauthorid">1610074-0</idno>
                  <affiliation ref="#struct-300666"/>
                </author>
                <author role="aut">
                  <persName>
                    <forename type="first">Onar</forename>
                    <surname>Vedat</surname>
                  </persName>
                  <idno type="idhal" notation="numeric">1178464</idno>
                  <idno type="halauthorid" notation="string">2761086-1178464</idno>
                  <idno type="ORCID">https://orcid.org/0000-0002-8359-243X</idno>
                  <idno type="RESEARCHERID">http://www.researcherid.com/rid/D-1582-2019</idno>
                  <idno type="RESEARCHERID">http://www.researcherid.com/rid/http://www.researcherid.com/rid/D-1582-2019</idno>
                  <affiliation ref="#struct-495165"/>
                </author>
                <author role="crp">
                  <persName>
                    <forename type="first">Ludovic</forename>
                    <surname>Orlando</surname>
                  </persName>
                  <email type="md5">7585ec3f47399a860c08466925c6c7e0</email>
                  <email type="domain">univ-tlse3.fr</email>
                  <idno type="idhal" notation="string">ludovicorlando</idno>
                  <idno type="idhal" notation="numeric">1083415</idno>
                  <idno type="halauthorid" notation="string">312103-1083415</idno>
                  <idno type="ORCID">https://orcid.org/0000-0003-3936-1850</idno>
                  <idno type="IDREF">https://www.idref.fr/080106951</idno>
                  <idno type="RESEARCHERID">http://www.researcherid.com/rid/A-8932-2013</idno>
                  <idno type="RESEARCHERID">http://www.researcherid.com/rid/http://www.researcherid.com/rid/A-8932-2013</idno>
                  <idno type="GOOGLE SCHOLAR">https://scholar.google.fr/citations?user=zSlox0oAAAAJ</idno>
                  <idno type="VIAF">https://viaf.org/viaf/66751689</idno>
                  <idno type="ISNI">http://isni.org/isni/0000000041962127</idno>
                  <affiliation ref="#struct-445683"/>
                  <affiliation ref="#struct-3549"/>
                </author>
              </analytic>
              <monogr>
                <idno type="halJournalId" status="VALID">15097</idno>
                <idno type="issn">0305-4403</idno>
                <idno type="eissn">1095-9238</idno>
                <title level="j">Journal of Archaeological Science</title>
                <imprint>
                  <publisher>Elsevier</publisher>
                  <biblScope unit="volume">78</biblScope>
                  <biblScope unit="pp">147-157</biblScope>
                  <date type="datePub">2017-02</date>
                </imprint>
              </monogr>
              <idno type="doi">10.1016/j.jas.2016.12.005</idno>
            </biblStruct>
          </sourceDesc>
          <profileDesc>
            <langUsage>
              <language ident="en">English</language>
            </langUsage>
            <textClass>
              <keywords scheme="author">
                <term xml:lang="en">Hinny</term>
                <term xml:lang="en">Mule</term>
                <term xml:lang="en">F1-hybrid</term>
                <term xml:lang="en">Equid</term>
                <term xml:lang="en">High-throughput sequencing</term>
                <term xml:lang="en">Ancient DNA</term>
              </keywords>
              <classCode scheme="halDomain" n="sdv.gen">Life Sciences [q-bio]/Genetics</classCode>
              <classCode scheme="halDomain" n="shs.archeo">Humanities and Social Sciences/Archaeology and Prehistory</classCode>
              <classCode scheme="halTypology" n="ART">Journal articles</classCode>
              <classCode scheme="halOldTypology" n="ART">Journal articles</classCode>
              <classCode scheme="halTreeTypology" n="ART">Journal articles</classCode>
            </textClass>
            <abstract xml:lang="en">
              <p>Horses, asses and zebras, can produce first-generation F1-hybrids, despite their striking karyotypic and phenotypic differences. Such F1-hybrids are mostly infertile, but often present characters of considerable interest to breeders. They were extremely valued in antiquity, and commonly represented in art and on coinage. However, hybrids appear relatively rarely in archaeological faunal assemblages, mostly because identification based on morphometric data alone is extremely difficult. Here, we developed a methodological framework that exploits high-throughput sequencing data retrieved from archaeological material to identify F1-equine hybrids. Our computational methodology is distributed in the open-source Zonkey pipeline, now part of PALEOMIX (https://github.com/MikkelSchubert/paleomix), together with full documentation and examples. Using both synthetic and real sequence datasets, from living and ancient F1-hybrids, we find that Zonkey shows high sensitivity and specificity, even with limited sequencing efforts. Zonkey is thus well suited to the identification of equine F1-hybrids in the archaeological record, even in cases where DNA preservation is limited. Zonkey can also help determine the sex of ancient animals, and allows species identification, which advantageously complements morphological data in cases where material is fragmentary and/or multiple candidate equine species coexisted in sympatry.</p>
            </abstract>
          </profileDesc>
        </biblFull>
      </listBibl>
    </body>
    <back>
      <listOrg type="structures">
        <org type="department" xml:id="struct-3549" status="VALID">
          <orgName>Geological Museum [Copenhagen]</orgName>
          <desc>
            <address>
              <addrLine>Olster Voldgade 5-7, Copenhagen K., Denmark</addrLine>
              <country key="DK"/>
            </address>
          </desc>
          <listRelation>
            <relation active="#struct-478580" type="direct"/>
            <relation active="#struct-579313" type="indirect"/>
            <relation active="#struct-460939" type="indirect"/>
          </listRelation>
        </org>
        <org type="laboratory" xml:id="struct-1225933" status="VALID">
          <orgName>BioArchéologie, Interactions Sociétés Environnements</orgName>
          <orgName type="acronym">BioArch</orgName>
          <date type="start">2025-01-01</date>
          <desc>
            <address>
              <addrLine>Muséum national d’Histoire naturelleCP 56 - 43 rue Buffon75005 Paris - France</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">https://archeozoo-archeobota.mnhn.fr/fr</ref>
          </desc>
          <listRelation>
            <relation active="#struct-7512" type="direct"/>
            <relation name="UMR7209" active="#struct-441569" type="direct"/>
          </listRelation>
        </org>
        <org type="laboratory" xml:id="struct-445683" status="OLD">
          <idno type="IdRef">238573346</idno>
          <idno type="RNSR">201119472X</idno>
          <idno type="ROR">https://ror.org/05w482q29</idno>
          <orgName>Anthropologie Moléculaire et Imagerie de Synthèse</orgName>
          <orgName type="acronym">AMIS</orgName>
          <date type="start">2009-01-01</date>
          <date type="end">2020-12-31</date>
          <desc>
            <address>
              <addrLine>37 Allées Jules Guesde31073 Toulouse</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://amis.cnrs.fr/</ref>
          </desc>
          <listRelation>
            <relation active="#struct-217752" type="direct"/>
            <relation active="#struct-443875" type="indirect"/>
            <relation name="UMR5288" active="#struct-441569" type="direct"/>
          </listRelation>
        </org>
        <org type="institution" xml:id="struct-302265" status="VALID">
          <idno type="ROR">https://ror.org/02f81g417</idno>
          <orgName>King Saud University [Riyadh]</orgName>
          <orgName type="acronym">KSU</orgName>
          <desc>
            <address>
              <addrLine>King Saud University, Riyadh 12372</addrLine>
              <country key="SA"/>
            </address>
            <ref type="url">http://ksu.edu.sa/en/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-300666" status="VALID">
          <idno type="ROR">https://ror.org/01ryk1543</idno>
          <orgName>University of Southampton</orgName>
          <desc>
            <address>
              <addrLine>University Road, Southampton SO17 1BJ</addrLine>
              <country key="GB"/>
            </address>
            <ref type="url">http://www.southampton.ac.uk/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-495165" status="VALID">
          <idno type="ROR">https://ror.org/03a5qrr21</idno>
          <orgName>Istanbul University</orgName>
          <date type="start">1453-05-30</date>
          <desc>
            <address>
              <addrLine>Istanbul University Main Campus34452 Beyazit/Fatih-Istanbul</addrLine>
              <country key="TR"/>
            </address>
            <ref type="url">https://www.istanbul.edu.tr/tr/_</ref>
          </desc>
        </org>
        <org type="laboratory" xml:id="struct-478580" status="VALID">
          <orgName>Natural History Museum of Denmark</orgName>
          <desc>
            <address>
              <addrLine>Universitetsparken 15DK-2100 Copenhagen Ø</addrLine>
              <country key="DK"/>
            </address>
            <ref type="url">https://snm.ku.dk/english/</ref>
          </desc>
          <listRelation>
            <relation active="#struct-579313" type="direct"/>
            <relation active="#struct-460939" type="indirect"/>
          </listRelation>
        </org>
        <org type="regrouplaboratory" xml:id="struct-579313" status="VALID">
          <orgName>Faculty of Science [Copenhagen]</orgName>
          <desc>
            <address>
              <addrLine>Bülowsvej 171870 Frederiksberg CCopenhagen</addrLine>
              <country key="DK"/>
            </address>
            <ref type="url">https://www.science.ku.dk/english/</ref>
          </desc>
          <listRelation>
            <relation active="#struct-460939" type="direct"/>
          </listRelation>
        </org>
        <org type="institution" xml:id="struct-460939" status="VALID">
          <idno type="IdRef">026458748</idno>
          <idno type="ROR">https://ror.org/035b05819</idno>
          <orgName>Københavns Universitet = University of Copenhagen = Université de Copenhague</orgName>
          <orgName type="acronym">UCPH</orgName>
          <date type="start">1479-01-01</date>
          <desc>
            <address>
              <addrLine>Nørregade 10, 1165 København, Danemark</addrLine>
              <country key="DK"/>
            </address>
            <ref type="url">http://www.ku.dk</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-7512" status="VALID">
          <idno type="IdRef">026394944</idno>
          <idno type="ROR">https://ror.org/03wkt5x30</idno>
          <orgName>Muséum national d'Histoire naturelle</orgName>
          <orgName type="acronym">MNHN</orgName>
          <desc>
            <address>
              <addrLine>57, rue Cuvier - 75231 Paris Cedex 05</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://www.mnhn.fr</ref>
          </desc>
        </org>
        <org type="regroupinstitution" xml:id="struct-441569" status="VALID">
          <idno type="IdRef">02636817X</idno>
          <idno type="ISNI">0000000122597504</idno>
          <idno type="ROR">https://ror.org/02feahw73</idno>
          <orgName>Centre National de la Recherche Scientifique</orgName>
          <orgName type="acronym">CNRS</orgName>
          <date type="start">1939-10-19</date>
          <desc>
            <address>
              <country key="FR"/>
            </address>
            <ref type="url">https://www.cnrs.fr/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-217752" status="OLD">
          <idno type="IdRef">026404672</idno>
          <idno type="ISNI">0000000121617331</idno>
          <idno type="ROR">https://ror.org/02v6kpv12</idno>
          <idno type="Wikidata">Q1273188</idno>
          <orgName>Université Toulouse III - Paul Sabatier</orgName>
          <orgName type="acronym">UT3</orgName>
          <date type="end">2025-01-01</date>
          <desc>
            <address>
              <addrLine>118 route de Narbonne - 31062 Toulouse</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://www.univ-tlse3.fr/</ref>
          </desc>
          <listRelation>
            <relation active="#struct-443875" type="direct"/>
          </listRelation>
        </org>
        <org type="regroupinstitution" xml:id="struct-443875" status="VALID">
          <idno type="ROR">https://ror.org/017tgbk05</idno>
          <orgName>Communauté d'universités et établissements de Toulouse</orgName>
          <orgName type="acronym">Comue de Toulouse</orgName>
          <desc>
            <address>
              <addrLine>41 Allée Jules Guesde, 31000 Toulouse</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">https://www.univ-toulouse.fr/</ref>
          </desc>
        </org>
      </listOrg>
    </back>
  </text>
</TEI>